109
Role of Hydropriming and Magneto-Priming in Developing Stress Tolerance
et al., 1998). The crucial technique of hydropriming is drum priming, which
comprises a drum enclosed with seed coupled with a boiler creating vapors
that condense and water droplets form in the drum (Warren & Bennett, 1997).
Many studies have stated improved stand establishment, seed vigor, and
yield under suboptimal or optimal conditions along with three to four-fold
increases in germination as compared to non-primed seeds. Some studies
explained improved seed germination by hydropriming in wheat under saline
conditions (Afzal et al., 2007; Harris et al., 2001; Nawaz et al., 2016; Roy &
Srivastava, 1999), chickpea (Harris et al., 1999; Kamithi et al., 2016; Kaur
et al., 2002), lentil (Ghassemi et al., 2008), safflower (Ashrafi & Razmjoo,
2010; Bastia et al., 1999), mountain rye (Ansari & Zadeh, 2012), water
melon (Sung & Chiu, 1995), pearl millet (Kumar et al., 2002). Hydropriming
was effective in enhancing vegetative and reproductive growth stages and
seedling emergence in maize (Dezfuli et al., 2008; Mohammadi et al., 2008;
Nagar et al., 1998). Several food crops coriander, wheat, Allium porrum and
in desert plants like cacti (Dubrovsky, 1996) showed synchronized early
germination after hydropriming. Similarly, improved seed germination was
observed in onion (Caseiro et al., 2004) cauliflower (Jisha et al., 2013),
and in mustard (Srivastava et al., 2010). Therefore, hydropriming provides
improved seedling emergence seed germination, and productivity of field
crops that can be explained by the rapid emergence of shoots and roots, and
better tolerance under adverse conditions (Lee-Suskoon et al., 1998).
4.3.1 SALT
Salt stress is one of the leading problems and major growth-limiting factors in
agriculture, with one-third of the world’s land affected by salinity (Flowers &
Colmer, 2008). Other factors such as nonsystemic irrigation, natural weath
ering, or intense agriculture also contribute to salt stress (Hasanuzzaman &
Fotopoulos, 2019). The excess build-up of salts reduces water potential in
plants, as consequence plants don’t get access to water (Munns et al., 2006).
Further reduction of water and nutrient uptake due to ion toxicity is observed
(Chinnusamy et al., 2005). The impact of salinity includes, reduced water
and nutrient uptake, altered rate of respiration, reduced rate of photosyn
thesis, lowered transpiration, ion toxicity, and affects stomatal conductance.
When a plant is exposed to salinity, firstly the osmotic (hyperosmotic) stress
happens due to excess sodium in the root zone, followed by ionic (hypertonic)
stress due to altered concentration of essential nutrients (Munns et al., 2006).
An increase in sodium levels reduces potassium ion influx and disturbs the